Research Article | | Peer-Reviewed

Assessment and Identification of Major Weeds on Maize (Zea mays) in East Shewa and West Arsi, Zones, Oromia

Received: 8 December 2025     Accepted: 20 December 2025     Published: 7 January 2026
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Abstract

Major cereals globally include wheat, maize, and rice, contributing 80% of production. In Ethiopia, key cereals are tef, maize, wheat, barley, sorghum, rice, and finger millet. Maize accounts for 27.02% of annual grain production, followed by tef at 17.29%. Maize yields the highest per unit area at 3.7 t/ha, followed by rice and wheat. Ethiopia's mean national wheat yield is 2600.75 kg/ha, below African and global averages. Yield-reducing factors include soil fertility decline, weeds, diseases, and insects. Weeds significantly constrain cereal crop production and are crucial for yield increase. Weeds compete for resources, negatively impacting crop yield and increasing production costs. Weeds contribute to 45% of annual agricultural product losses globally. Crop yield losses due to weeds in Africa range from 50% to 100% for various crops. Limited studies exist on common weeds affecting Maize at the West Arsi and East shewa. The weed survey was conducted in East Shewa and West Arsi zones, Oromia. Assessment took place during the main cropping seasons of 2021-2023. The survey covered seven districts: ATJK, Dugda, Bora, Lume, Negelle Arsi, Shashemanne, and Kofele. Maize fields in 37 kebeles and 107 fields were included in the survey. Key parameters such as density, frequency, relative frequency and similarity index have been analyzed for each crop. Generally, for most crops and districts, annual broad leaf weeds dominated over grass and sedge type of weeds. The most dominant family that contained the highest number of weed species for all crops and fields assessed was Asteracae followed by Poaceace and Amaranthaceae families. In most cases weeds of a crop were similar across the various districts as indicated by similarity index. The frequency of individual weed species in maize fields ranged from 1% up to 91% while the dominance value ranged from 0.71% up to 21.92%. The most frequent and dominant weeds were Galensoga palviflora and Argemone mexicana L., Maize crop.

Published in American Journal of Plant Biology (Volume 11, Issue 1)
DOI 10.11648/j.ajpb.20261101.11
Page(s) 1-7
Creative Commons

This is an Open Access article, distributed under the terms of the Creative Commons Attribution 4.0 International License (http://creativecommons.org/licenses/by/4.0/), which permits unrestricted use, distribution and reproduction in any medium or format, provided the original work is properly cited.

Copyright

Copyright © The Author(s), 2026. Published by Science Publishing Group

Keywords

Maize, Assessment, Major Weeds, Galensoga Palviflora

1. Introduction
1.1. Background and Justification
Wheat, maize and rice contribute about 80% of the global cereal production. The major cereals grown in Ethiopia include tef (Eragrostistef), maize (Zea mays L.), bread wheat (Triticum aestivum L.), durum wheat (Triticum durum Desf.), barley (Hordeum vulgare L.), sorghum (Sorghum biclor L.), rice (Oryza sativa L.), and (Eleusine corocana L.) finger millet . The highest proportion of annual grain production of the country comes from maize which makes up to 27.02% followed by teff (17.29%) of the total produce. Maize also gives the highest yield per unit area (3.7 t/ha), followed by rice (2.8 t/ha), wheat (2.6 t/ha), sorghum (2.5 t/ha), barley (2.1 t/ha), and tef (1.7 t/ha), respectively .
Despite its importance in Ethiopia, the mean national yield is 2600.75kg/ha-1which is 12% below the mean yield of Africa and 24% below the global mean yield of wheat . Yield reducing factors in cereal crops are soil fertility decline, weeds, disease, and insects. Weeds are one of the major constraints of cereal crops production and weed control is the key factor in increasing yield . Weeds are undesirable plants, which infest different crops and inflict negative effect on crop yield either competition for water or nutrients or space or light . There are innumerable reports on the inhibitory effects of weeds on crop plants . Research indicates that over 80 weed species, predominantly broad-leaved, severely impact maize productivity, causing yield losses ranging from 35% to 87.5% due to competition and toxicity . Effective identification and management strategies are crucial for mitigating these losses. Although most farmers are less concerned about the negative impact that weeds impose on their crop, study results indicate that weeds share up to 45% of the total annual losses of agricultural products .
Weeds surveys are commonly used to characterize weed populations in cropping systems . Research indicated that, worldwide, over 10% of agricultural production is lost as a result of crop weed competition for the resources light, water and nutrients . According to when weeds are left uncontrolled, yield losses range from 20-100%, depending upon the crop and its environment. The author reported loss estimates of 5% in developed countries, 10% in the less developed countries and 25% in the least developed countries .
Therefore, to develop an effective weed management strategy, weed survey is necessary to address the current weed problems on major cereals crops. In addition, survey information is important in developing target-oriented research and control measure. So far, insufficient study has been conducted regarding the occurrence and distribution of common weeds associated with maize in central rift valley Ethiopia.
1.2. Objective
To identify and identify common weeds associated with Maize at a major production area of West Arsi and East Shewa.
2. Materials and Methods
2.1. Description of the Study Area
The weed survey was conducted in East Shewa and West Arsi zones of Oromia Regional state during main cropping seasons of 2021-2023. The weed assessment survey was conducted in the seven Districts of ATJK, Dugda, Bora, Lume, Negelle Arsi, Shashemanne, and Kofele. The weed survey was conducted to assess Density, Frequency, Relative Density, Relative Frequency and Similarity index of weeds. The geographical locations of the surveyed areas were located in the rage of latitude and longitude of 8° 34' 59.99" N and 39° 09' 60.00" E and 7° 09' 60.00" N and 38° 49' 59.99" E respectively (Figure 1).
Figure 1. Map of Survey Site.
2.2. Field Survey
Table 1. Characteristic Features of Surveyed Maize Fields in Two Zones of Study Area.

ZONE

Districs

Altitude (ab.ms.l)

no of field assesed

East Shewa

ATJK

1626-1841

13

Dudga

1633-1670

16

Bora

1590-1680

15

Lume

1586-1920

14

West Arsi

Negelle Arsi

1604-1967

21

Shashemanne

2168-2455

14

Kofele

2398-2646

14

Over all mean

1586-2646

107

no of kebele

37

M. A. S. L. = Meter Above Sea Level
The survey was conducted at the Maize fields in 37 kebeles and 107 fields in the seven Districts of the two zones. Purposive sampling technique was applied to select Districts. Purposive sampling technique was applied to select Districts. Kebele were Randomly selected from each Districts based on the potential production of the Maize. Consecutive sample sites for the same crop were 5 km apart depending on the topography and the relative importance of the crop within each location. Weed assessment was made along the two diagonals (in an “X” pattern) of the field from three points 1m × 1m (1m2) quadrates for maize. Frequency (F), Density (D) and Similarity Index (SI) were computed for each species of weeds using the method of Thomas . The collected weed data were combined and summarized. In each field, weeds specie and their numbers within the quadrates were counted and recorded.
Farmers were interviewed using pre-structured questionaries record information on farmers’ practices such as: - the cereal crop and management practices, variety/ies grown whether local or improved, previous crop (cereals, pulses or vegetables), planting date (sowing time), crop density, altitude, fertilizer type and rate, soil type, growth stage, disease type observed and herbicides used were collected as to the survey format.
2.3. Data Analysis
Density, Relative Density, Frequency, Relative Frequency and Similarity index were calculated by the following formula. The collected weed data were combined and summarized using MS Excel and Minitab (17.0) version software.
Density (D) =Total No of individuals of a species in all quadrantTotal No of quadran used 
Frequency (F)=No of quadrantes in which a given speceis ocuursTotal No of quadrant used
Relative density (RD) =Densityof a given species Rotal density for all spcies X100
Relative frequency (RF)=Frequency of a given species Total frequency for all species X100
Summed dominant ratio (SDR) =Relative density Relative frequency X100
Similarity Index (SI) = Epg/(Epg + Epa + Epb) X 100
Where; SI = Similarity index, Epg = number of species found in both locations, Epa = number of species found only in location I. Epb = number of species found only in locations II.
3. Result and Discussion
3.1. Diversity of Weeds
Table 2. Number of Weed Families and Number of Species They Comprise in the Maize Fields.

No.

Family

No. of species

No.

Family

No. of species

1

Asteraceae

9

11

Commelinaceae

1

2

Poaceae

7

12

Compositae

1

3

Amaranthaceae

4

13

Eragrostidae

1

4

Lamiaceae

3

14

Malvaceae

1

5

Solanaceae

3

15

Oxalidaceae

1

6

Brassicaceae

2

16

Panicae

1

7

Convolvulaceae

2

17

Papaveraceae

1

8

Cyperaceae

2

18

Primulaceae

1

9

Euphorbiaceae

2

19

Protulaceae

1

10

Polygonaceae

2

20

Rubiaceae

1

Total

46

In the surveyed Maize fields, 46 weed species that belonged to 20 families were recorded of which 46 species (72.34%) were annuals, 11 weed species (23.40%) were perennials and 2 species (4.25%) were Biennials. About 78.26% were Broad leaf; 17.4% were grass type and 4.35 were sedge types of weed species.
3.2. Weed Flora of Maize Fields
The frequency of occurrence of individual weed species ranged from 1%-91% (Table 3). Dominant weeds were those species which occurred in relatively greater number than the others species. Ten of weed species recorded in Maize fields i.e Cynodon dactlyon, Digitaria diagonalis, Cyprus esculentus, Bidense Pilosa, Gozotia abisinica Argemone mexicana L., Amaranthus spinosus, Commelina benghalensis, Nicandra physlodes, Galensoga palviflora were widely distributed with higher than 35% frequency while 10 weed species have less than 30% frequency value. The species have highest frequency 91% was Galensoga palviflora followed by Nicandra physlodes, Argemone mexicana L, Amaranthus spinosus and Commelina benghalensis which scores 62%, 58%, 56% and 52% respectively. This indicates broad leaf is dominated over grass weed in the Maize field similar study with which indicates dominant species broad-leaved weeds such as Ipomea eriocarpa and Setaria viridis are prevalent in maize fields.
Table 3. Description of Density, Frequency, Relative Density, and Relative Frequency of Weed in Maize Fields.

Botaical Name

family

Catagories

Life cycle

Density

Frequ ency

RD

FR

SD

Agerantum conyoides

Asteraceae

Broad Leaf

Annual

2.18

0.11

1.38

1.24

111.50

Amaranthus hybridus

Amaranthaceae

Broad Leaf

Annual

1.53

0.05

0.97

0.56

172.39

Amaranthus retroflexus

Amaranthaceae

Broad Leaf

Biennial

1.33

0.01

0.84

0.11

747.65

Amaranthus spinosus

Amaranthaceae

Broad Leaf

Annual

2.07

0.56

1.31

6.19

21.12

Anagallis arvensis

Primulaceae

Broad Leaf

Annual

1.90

0.13

1.20

1.46

82.05

Argemone mexicana L

Papaveraceae

Broad Leaf

Annual

4.20

0.52

2.65

5.74

46.25

Avena fatua

Poaceae

Grassy

Annual

2.76

0.15

1.74

1.69

103.27

Bidense pilosa

Asteraceae

Broad Leaf

Annual

2.07

0.38

1.31

4.16

31.48

Brassica juncea

Brassicaceae

Broad Leaf

Annual

0.95

0.14

0.60

1.58

38.24

Chenopodium album

Amaranthaceae

Broad Leaf

Annual

2.36

0.23

1.49

2.59

57.74

Commelina benghalensis

Commelinaceae

Broad Leaf

Annual

2.29

0.58

1.45

6.41

22.55

Convolvulus

Convolvulaceae

Broad Leaf

Pennerial

0.78

0.06

0.49

0.68

72.87

Cyanthillium cinereum

Brassicaceae

Broad Leaf

Biennial

4.00

0.01

2.53

0.11

2248.57

Cynodon dactlyon

Poaceae

Grassy

Pennerial

3.57

0.35

2.26

3.83

59.00

Cyprus esculentus

Cyperaceae

Sedge

Pennerial

7.10

0.37

4.49

4.05

110.90

Cyprus rotundus

Cyperaceae

Sedge

Pennerial

4.52

0.07

2.86

0.79

363.29

Datura stramonium

Solanaceae

Broad Leaf

Annual

1.17

0.18

0.74

2.03

36.44

Digitaria diagonalis

Poaceae

Grassy

Pennerial

4.30

0.36

2.72

3.94

69.14

Echinochloa esculenta

Poaceae

Grassy

Pennerial

3.87

0.05

2.45

0.56

434.72

Eleusine indica

Poaceae

Grassy

Pennerial

2.70

0.18

1.71

2.03

84.44

Eragrostis cilianen

Eragrostidae

Grassy

Annual

3.28

0.20

2.08

2.25

92.29

Euphorbia esula

Euphorbiaceae

Broad Leaf

Pennerial

1.20

0.10

0.76

1.13

67.46

Euphorbia hirta

Euphorbiaceae

Broad Leaf

Annual

21.92

0.04

13.86

0.45

3080.07

Galensoga palviflora

Asteraceae

Broad Leaf

Annual

8.63

0.91

5.46

10.01

54.48

Galium spurium

Rubiaceae

Broad Leaf

Annual

2.73

0.05

1.73

0.56

307.30

Gozotia abisinica

Asteraceae

Broad Leaf

Annual

2.37

0.39

1.50

4.28

35.04

Ipomoea indica

Convolvulaceae

Broad Leaf

Annual

0.71

0.07

0.45

0.79

57.36

Leucas aspera

Lamiaceae

Broad Leaf

Annual

1.74

0.09

1.10

1.01

108.73

Malva parviflora

Malvaceae

Broad Leaf

Annual

2.80

0.05

1.77

0.56

314.80

Nicandra physlodes

Solanaceae

Broad Leaf

Annual

2.88

0.62

1.82

6.86

26.54

Ocimum lamiifolium

Lamiaceae

Broad Leaf

Annual

1.11

0.03

0.70

0.34

208.20

Oxalis acetosella

Oxalidaceae

Broad Leaf

Pennerial

4.29

0.07

2.71

0.79

344.17

Oxygonum sinuatum Meisn.

Polygonaceae

Broad Leaf

Annual

1.53

0.19

0.97

2.14

45.16

Parthiunium Hysterophoros

Asteraceae

Broad Leaf

Annual

1.88

0.20

1.19

2.25

52.94

Pennisetum setaceum

Poaceae

Grassy

Pennerial

3.95

0.21

2.50

2.36

105.80

Polygonum arenastrum

Polygonaceae

Broad Leaf

Annual

8.42

0.08

5.32

0.90

591.42

Portulaca oleraceus

Protulaceae

Broad Leaf

Annual

1.33

0.06

0.84

0.68

124.92

salvia officinalis

Lamiaceae

Broad Leaf

Pennerial

2.53

0.15

1.60

1.69

94.94

Setaria pumila

Poaceae

Broad Leaf

Annual

3.22

0.09

2.04

1.01

201.26

Setria verticelata

Panicae

Grassy

Annual

3.06

0.27

1.94

2.93

66.25

Silybum marianum

Asteraceae

Broad Leaf

Biennial

2.78

0.03

1.76

0.34

520.29

Solanum nigrum

Solanaceae

Broad Leaf

Annual

2.13

0.05

1.35

0.56

239.92

Sonchus oleraceus

Compositae

Broad Leaf

Annual

0.83

0.02

0.53

0.23

234.23

Tagetes minuta

Asteraceae

Broad Leaf

Annual

8.60

0.05

5.44

0.56

966.89

Xanthium spinosum

Asteraceae

Broad Leaf

Annual

0.88

0.08

0.55

0.90

61.48

Xanthium strumarium

Asteraceae

Broad Leaf

Annual

1.61

0.29

1.02

3.15

32.27

3.3. Weed Similarity Index
Similarity index (community index) is the similarity of plant species composition among different districts. When the percent of the Weed Similarity Index (WSI) were greater than 60% thus require /similar weed management options and vice versa. The survey result showed that similarity index value between ATJK and Dugda, ATJK and Lume and ATJK and Negelle Arsi than 60% (Table 4); it can be concluded that the locations exhibited similar weed community and thus require similar management options. While ATJK and Bora, ATJK and Shashemanne. Kofele were less than 60%. Also, Dugda and Bora and Lume were greater than 60. While Dugda and Negelle Arsi, Shashemanne and Kofele were performed less than 60% weed similarity index. Bora and Lume showed 69.23%. while Bora and Negelle Arsi, Shashemanne and Kofele were less 60%. Even if Bora and Shashemanne were huge difference (29.89%) from rest of others. Lume and Negelle Arsi were (64.90% WSI) while Lume and Shashemanne and Kofele were Less than 60%. Negelle Arsi and Shashemanne were showed 45.24% and Negelle arsi and Kofele were 60%. Finally, the weed similarity index (WSI) between Shashemanne and Kofele recorded during survey was 39.39%. This due to agro-ecological location different and different weed management practices similarly reported .
Table 4. Number of Weed Families Identified and Number of Species They Comprise in Maize Fields.

District

ATJK

Dugda

Bora

Lume

Negelle A

Shashe

Kofele

ATJK

100.00%

69.23%

58.13%

70.20%

61.90%

38.09%

50.00%

Dugda

100.00%

63.41%

70.20%

53.49%

41.46%

46.20%

Bora

100.00%

69.23%

54.54%

29.89%

46.15%

Lume

100.00%

64.90%

50.00%

47.30%

Negelle A

100.00%

45.24%

60.00%

Shashe

100.00%

39.39%

Kofele

100.00%

4. Conclusion
In the current study, a total of 107 fields were surveyed for weed flora and fauna of Maize crop, and different weed families and species were identified in the East Shewa and West Arsi zones. The importance of each species was determined by calculating the Frequency, dominant, Density and similarity index values. Generally, annual broad weed leaves dominated over grass and sedge types for Maize. The most dominant families according to frequency and number of weed species were Asteraceae, Poaceace, Solanaceae and Amaranthaceae. The most frequent and dominant weed species consisted of Galensoga Palviflora, Argemen Mexicana and Nicandra physlodas for Maize crop fields.
The current study has documented important weeds of Maize in representative and potential Agro-ecologies of the respective crops. As the weeds recorded were described in detail – by families, species and frequency, this information can be useful to prioritize weed management research and management strategies to pursue in the future for the various crops and districts.
5. Recommendation
5.1. Districts with Similar Weed Species for the Maize Fields
Districts with a WSI greater than 60% (e.g., ATJK and Dugda, ATJK and Lume, Dugda and Bora, etc.) should implement similar weed management practices due to their similar weed species compositions.
5.2. Districts with Diverse Weed Species
Districts with a WSI less than 60% (e.g., ATJK and Shashemanne, Dugda and Shashemanne, Bora and Shashemanne, etc.) require tailored weed management strategies to address their unique weed communities.
Abbreviations

ATJK

Adami Tulu Jido Combolcha

MSL

Above Sea Level

Conflicts of Interest
The authors declare no conflicts of interest.
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    Tesfaye, G., Begna, F., Longle, A. (2026). Assessment and Identification of Major Weeds on Maize (Zea mays) in East Shewa and West Arsi, Zones, Oromia. American Journal of Plant Biology, 11(1), 1-7. https://doi.org/10.11648/j.ajpb.20261101.11

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    Tesfaye, G.; Begna, F.; Longle, A. Assessment and Identification of Major Weeds on Maize (Zea mays) in East Shewa and West Arsi, Zones, Oromia. Am. J. Plant Biol. 2026, 11(1), 1-7. doi: 10.11648/j.ajpb.20261101.11

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    AMA Style

    Tesfaye G, Begna F, Longle A. Assessment and Identification of Major Weeds on Maize (Zea mays) in East Shewa and West Arsi, Zones, Oromia. Am J Plant Biol. 2026;11(1):1-7. doi: 10.11648/j.ajpb.20261101.11

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  • @article{10.11648/j.ajpb.20261101.11,
      author = {Gobena Tesfaye and Feyisa Begna and Adisu Longle},
      title = {Assessment and Identification of Major Weeds on Maize (Zea mays) in East Shewa and West Arsi, Zones, Oromia},
      journal = {American Journal of Plant Biology},
      volume = {11},
      number = {1},
      pages = {1-7},
      doi = {10.11648/j.ajpb.20261101.11},
      url = {https://doi.org/10.11648/j.ajpb.20261101.11},
      eprint = {https://article.sciencepublishinggroup.com/pdf/10.11648.j.ajpb.20261101.11},
      abstract = {Major cereals globally include wheat, maize, and rice, contributing 80% of production. In Ethiopia, key cereals are tef, maize, wheat, barley, sorghum, rice, and finger millet. Maize accounts for 27.02% of annual grain production, followed by tef at 17.29%. Maize yields the highest per unit area at 3.7 t/ha, followed by rice and wheat. Ethiopia's mean national wheat yield is 2600.75 kg/ha, below African and global averages. Yield-reducing factors include soil fertility decline, weeds, diseases, and insects. Weeds significantly constrain cereal crop production and are crucial for yield increase. Weeds compete for resources, negatively impacting crop yield and increasing production costs. Weeds contribute to 45% of annual agricultural product losses globally. Crop yield losses due to weeds in Africa range from 50% to 100% for various crops. Limited studies exist on common weeds affecting Maize at the West Arsi and East shewa. The weed survey was conducted in East Shewa and West Arsi zones, Oromia. Assessment took place during the main cropping seasons of 2021-2023. The survey covered seven districts: ATJK, Dugda, Bora, Lume, Negelle Arsi, Shashemanne, and Kofele. Maize fields in 37 kebeles and 107 fields were included in the survey. Key parameters such as density, frequency, relative frequency and similarity index have been analyzed for each crop. Generally, for most crops and districts, annual broad leaf weeds dominated over grass and sedge type of weeds. The most dominant family that contained the highest number of weed species for all crops and fields assessed was Asteracae followed by Poaceace and Amaranthaceae families. In most cases weeds of a crop were similar across the various districts as indicated by similarity index. The frequency of individual weed species in maize fields ranged from 1% up to 91% while the dominance value ranged from 0.71% up to 21.92%. The most frequent and dominant weeds were Galensoga palviflora and Argemone mexicana L., Maize crop.},
     year = {2026}
    }
    

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  • TY  - JOUR
    T1  - Assessment and Identification of Major Weeds on Maize (Zea mays) in East Shewa and West Arsi, Zones, Oromia
    AU  - Gobena Tesfaye
    AU  - Feyisa Begna
    AU  - Adisu Longle
    Y1  - 2026/01/07
    PY  - 2026
    N1  - https://doi.org/10.11648/j.ajpb.20261101.11
    DO  - 10.11648/j.ajpb.20261101.11
    T2  - American Journal of Plant Biology
    JF  - American Journal of Plant Biology
    JO  - American Journal of Plant Biology
    SP  - 1
    EP  - 7
    PB  - Science Publishing Group
    SN  - 2578-8337
    UR  - https://doi.org/10.11648/j.ajpb.20261101.11
    AB  - Major cereals globally include wheat, maize, and rice, contributing 80% of production. In Ethiopia, key cereals are tef, maize, wheat, barley, sorghum, rice, and finger millet. Maize accounts for 27.02% of annual grain production, followed by tef at 17.29%. Maize yields the highest per unit area at 3.7 t/ha, followed by rice and wheat. Ethiopia's mean national wheat yield is 2600.75 kg/ha, below African and global averages. Yield-reducing factors include soil fertility decline, weeds, diseases, and insects. Weeds significantly constrain cereal crop production and are crucial for yield increase. Weeds compete for resources, negatively impacting crop yield and increasing production costs. Weeds contribute to 45% of annual agricultural product losses globally. Crop yield losses due to weeds in Africa range from 50% to 100% for various crops. Limited studies exist on common weeds affecting Maize at the West Arsi and East shewa. The weed survey was conducted in East Shewa and West Arsi zones, Oromia. Assessment took place during the main cropping seasons of 2021-2023. The survey covered seven districts: ATJK, Dugda, Bora, Lume, Negelle Arsi, Shashemanne, and Kofele. Maize fields in 37 kebeles and 107 fields were included in the survey. Key parameters such as density, frequency, relative frequency and similarity index have been analyzed for each crop. Generally, for most crops and districts, annual broad leaf weeds dominated over grass and sedge type of weeds. The most dominant family that contained the highest number of weed species for all crops and fields assessed was Asteracae followed by Poaceace and Amaranthaceae families. In most cases weeds of a crop were similar across the various districts as indicated by similarity index. The frequency of individual weed species in maize fields ranged from 1% up to 91% while the dominance value ranged from 0.71% up to 21.92%. The most frequent and dominant weeds were Galensoga palviflora and Argemone mexicana L., Maize crop.
    VL  - 11
    IS  - 1
    ER  - 

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Author Information
  • Oromia Agricultural Research Institute, Adami Tullu Agricultural Research Center, Ziway, Ethiopia

  • Oromia Agricultural Research Institute, Adami Tullu Agricultural Research Center, Ziway, Ethiopia

  • Oromia Agricultural Research Institute, Adami Tullu Agricultural Research Center, Ziway, Ethiopia

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    1. 1. Introduction
    2. 2. Materials and Methods
    3. 3. Result and Discussion
    4. 4. Conclusion
    5. 5. Recommendation
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